Bold = lab members/students

Accepted/In press

  • Velasco, J.A., Villalobos, F., Diniz-Filho, J.A.F., Poe, S., Flores-Villela, O. Macroecology and macroevolution of body size in Anolis lizards. Ecography (In press) doi: 10.1111/ecog.04583
  • González-Tokman, D., Córdoba-Aguilar, A., Dáttilo, W., Lira-Noriega, A., Sánchez-Guillén, R.A., Villalobos, F.. Insect responses to heat: physiological mechanisms, evolution and ecological implications in a warming world. Biological Reviews (In press) doi: 10.1111/brv.12588
  • Nori, J., Loyola, R., Villalobos, F.. Priority areas for conservation of and research focused on terrestrial vertebrates. Conservation Biology (In Press) doi:10.1111/cobi.13476
  • Alves, D.M.C.C.,Eduardo, A.A., da Silva Oliveira, E. V., Villalobos, F., Dobrovolski, R., Pereira, T. C., Ribeiro, A., Stropp, J., Rodrigues, J.F., Diniz-Filho, J.A.F., Gouveia, S.F. Unveiling geographical gradients of species richness from scant occurrence data. Global Ecology and Biogeography (In Press) doi: 10.1111/geb.13055
  • Sanchez-Barradas, A., Villalobos, F..Species geographical co-occurrence and the effect of Grinnellian and Eltonian niche partitioning: the case of a Neotropical felid assemblage. Ecological Research (In press) doi: 10.1111/1440-1703.12070
  • Dáttilo, W.,Vázquez-Bolaños, M.,...Villalobos, F., et al. MEXICO ANTS: incidence and abundance along the Nearctic‐Neotropical interface. Ecology (In press) doi: 10.1002/ecy.2944


  • Kusumoto, B., Villalobos, F., Shiono, T., Kubota, Y. 2019. Reconciling Darwin’s naturalization and pre‐adaptation hypotheses: An inference from phylogenetic fields of exotic plants in Japan. Journal of Biogeography 46, 2597-2608. doi: 10.1111/jbi.13683
  • Carvajal-Quintero, J.D., Villalobos, F., Oberdorff, T., Grenouillet, G., Brosse, S., Hugueny, B., Jézéquel, C., Tedesco, P. 2019. Drainage network position and historical connectivity explain global patterns in freshwater fishes range size. Proceedings of the National Academy of Sciences 116, 13434-13439. doi:10.1073/pnas.1902484116
  • Diniz-Filho, J.A.F., Kelly S Souza, Luis M Bini, Rafael Loyola, Ricardo Dobrovolski, Fabricio Rodrigues, Matheus de S Lima-Ribeiro, Levi C Terribile, Thiago F Rangel, Igor Bione, Roniel Freitas, Iberê F Machado, Tainá Rocha, Maria L Lorini, Mariana M Vale, Carlos A Navas, Natan M Maciel, Fabricio Villalobos, Miguel A Olalla-Tarraga, Sidney Gouveia. 2019. A macroecological approach to evolutionary rescue and adaptation to climate change. Ecography 42, 1124-1141. doi: 10.1111/ecog.04264
  • Olalla-Tárraga, M.A., Amado, T.F., Bini, L.M., Martínez, P.A., Morales-Castilla, I., Torres-Romero, E.J., Villalobos, F.. 2019. Biological traits, phylogeny and human impacts effects on the geographic range size of passerines (Order Passeriformes) worldwide. Global Ecology and Biogeography 28, 1183-1194. doi: 10.1111/geb.12924
  • Velazco, S.J.E., Villalobos, F., Galvão, F., de Marco Jr., P. 2019. A dark scenario for Cerrado plant species: Effects of future climate, land use and protected areas ineffectiveness. Diversity and Distributions 25, 660-673. doi: 10.1111/ddi.12886
  • Rodrigues, J.F.M., Villalobos, F., Iverson, J.B., Diniz-Filho, J.A.F. 2019. Climatic niche evolution in turtles is characterized by phylogenetic conservatism for both aquatic and terrestrial species. Journal of Evolutionary Biology 32, 66-75. doi: 10.1111/jeb.13395
  • Pinto-Ledezma, J.N., Jahn, A.E., Cueto, V.R., Diniz-Filho, J.A.F., Villalobos, F. 2019. Drives of phylogenetic assemblage structure of the Furnariides, a widespread clade of lowland Neotropical birds. The American Naturalist 193, E41-E56. doi: 10.1086/700696


  • Nori, J., Villalobos, F., Loyola, R. 2018. Global priority areas for amphibian research. Journal of Biogeography 45, 2588-2594. doi: 10.1111/jbi.13435
  • Villalobos, F., Lira-Noriega, A., Rodríguez, P. 2018. Is species geographic overlap constant across latitude? A homage to E. H. Rapoport. [Special Issue on Macroecology and Geographical Ecology]. Oecologia Australis 22(2), 144-155. doi: 10.4257/oeco.2018.2202.04
  • Velasco, J.A., Villalobos, F., Diniz-Filho, J.A.F., Algar, A.C., Flores-Villela, O., Köhler, G., Poe, S., and Martínez-Meyer, E. 2018. Climatic and evolutionary factors shaping geographical gradients of species richness in Anolis lizards. Biological Journal of the Linnean Society 123 (3), 615-627. DOI: 10.1093/biolinnean/blx160
  • Alves, D.M.C.C., Diniz-Filho, J.A.F., Souza, K., Gouveia, S.F., Villalobos, F.. 2018. Geographic variation in the relationship between large-scale environmental determinants and bat species richness. Basic and Applied Ecology 27, 1-8. DOI: 10.1016/j.baae.2017.12.002
  • Bennet, J.M., Algar, A.C., Araújo, M.B., Calosi, P., Clusella-Trullas, S., Hawkins, B.A., Keith, S., Kühn, I., Martínez, B., Rahbek, C., Rodríguez, L., Singer, A., Sunday, J., Villalobos, F., Olalla-Tárraga, M.A. and Morales-Castilla, I. 2018. GlobTherm: a global database on thermal tolerances for aquatic and terrestrial organisms. Nature Scientific Data 5, 180022. DOI: 10.1038/sdata.2018.22


  • Rodríguez, P., Villalobos, F., Sánchez-Barradas, A., and Correa-Cano, .M.E. 2017. La macroecología en México: historia, avances y perspectivas. Revista Mexicana de Biodiversidad 88 (1), 52-64
  • Barreto, E., Pinto-Ledezma, J., Gomes de Freitas, C., Villalobos, F.,Collevati, R.G., and Maciel, N.M. 2017. Evolution of anuran foam nest: trait conservatism and lineage diversification. Biological Journal of the Linnean Society 122 (4), 814-823
  • Velazco, S.J.E., Galvão, F., Villalobos, F., de Marco Jr., P. 2017. Using worldwide edaphic data to model plant species niches: an assessment at a continental extent. PLoS ONE 12 (10), e0186025
  • Peixoto, F.P., Villalobos, F., and Cianciaruso, M. 2017. Phylogenetic conservatism of climatic niche in bats. Global Ecology and Biogeography 26, 1055-1065. DOI:10.1111/geb.12618
  • Barbosa, C., Otalora, J.M., Giehl, E.L.H., Villalobos, F., Loyola, R., Tessarolo, G., Machado, N. and Castellani T.T. 2017. Changes in the realised niche of the invasive succulent CAM plant Furcraea foetida. Austral Ecology 42, 643-654. doi:10.1111/aec.12483
  • Alves, D.M.C.C., Diniz-Filho, J.A.F., Villalobos, F.. 2017. Geographical diversification and the effect of model and data inadequacies: the bat diversity gradient as a case study. Biological Journal of the Linnean Society 121, 894-906. doi: 10.1093/biolinnean/blx030
  • Pinto-Ledezma J.N., Simon, L.M., Diniz-Filho, J.A.F., and Villalobos, F. 2017. The geographic diversification of Furnariides: the role of forest versus open habitats in driving species richness gradients. Journal of Biogeography 44, 1683-1693. doi:10.1111/jbi.12939
  • Hawkins, B.A., Leroy, B., Rodríguez, M.A., Singer, A., Vilela, B., Villalobos, F., Wang, X., and Zeleny, D. 2017. Structural bias in aggregated species-level variables driven by repeated species co-occurrences: a pervasive problem in community and assemblage data. Journal of Biogeography 44, 1199-1211. doi: 10.1111/jbi.12953
  • Peixoto, F.P.Villalobos, F., Melo, A.S., Diniz-Filho, J.A.F., Loyola, R., Rangel, T.F. and Cianciaruso, M. 2017. Geographical patterns of phylogenetic beta diversity components in terrestrial mammals. Global Ecology and Biogeography 26, 573-583. doi: 10.1111/geb.12561
  • Alves, D.M.C.C., Diniz-Filho, J.A.F., and Villalobos, F. 2017. Integrating Selection, Niche and Diversification into a Hierarchical Conceptual Framework. Organisms Diversity and Evolution 17, 1-10. doi:10.1007/s13127-016-0299-x
  • Villalobos, F., Cianciaruso, M.V., Olalla-Tárraga, M.A., Rangel, T.F., and Diniz-Filho, J.A.F. 2017. Global patterns of mammalian co-occurrence: phylogenetic and body size structure within species ranges. Journal of Biogeography 44, 136-146. doi:10.1111/jbi.12826
  • Olalla-Tárraga, M.A., González-Suárez, M., Bernardo-Madrid, R., Revilla, E., and Villalobos, F. 2017. Contrasting evidence of phylogenetic trophic niche conservatism in mammals worldwide. Journal of Biogeography 44, 99-110. doi:10.1111/jbi.12823
  • Villalobos, F., Olalla-Tárraga, M.A., Vieira, C.M., Mazzei, N.D., and Bini, L.M. 2017. Spatial dimension of body size evolution in Pterosauria: Bergmann's rule does not drive Cope's rule. Evolutionary Ecology Research 18, 169-186.


  • Villalobos, F., Carotenuto F., Raia, P., and Diniz-Filho, J.A.F. 2016. Phylogenetic fields through time: temporal dynamics of geographic co-occurrence and phylogenetic structure within species ranges. Philosophical Transactions of the Royal Society B 371, 20150220 DOI: 10.1098/rstb.2015.0220
  • Moura, M.R., Villalobos, F., Costa, G.C., and Garcia, P.C.A. Disentangling the role of climate, topography and vegetation in species richness gradients. PLoS ONE 11(3): e0152468
  • Morales, A., Villalobos, F., Velazco, P.M., Simmons, N., and Piñero, D. 2016. Environmental niche may drive genetic and morphometric structure in a widespread bat. Journal of Biogeography 43, 1057:1068, doi:10.1111/jbi.12666


  • Grandez-Rios, J.M., Bergamini, L.L., Araújo, W.S., Villalobos, F., and Almeida-Neto, M. 2015. The effect of host phylogenetic isolation on species richness, composition and specialization of insect herbivores: a comparison between native and exotic plants. PLOS ONE doi:10.1371/journal.pone.0138031
  • Diniz-Filho, J.A.F., Alves, D.M.C.C, Villalobos, F., Sakamoto, M., Brusatte, S., Bini, L.M. 2015. Phylogenetic eigenvectors and non-stationarity in the evolution of theropod dinosaur skulls. Journal of Evolutionary Biology 28, 1410:1416. doi: 10.1111/jeb.12660
  • Diniz-Filho, J.A.F., Villalobos, F. and Bini, L.M. 2015. The best of both worlds: phylogenetic eigenvector regression and mapping. Genetics and Molecular Biology 38, 396-400.
  • Vilela, B. and Villalobos, F. 2015. letsR: a new R package for data handling and analysis in macroecology. Methods in Ecology and Evolution. DOI: 10.1111/2041-210X.12401. The package is available from the CRAN repository of R packages ( and the package's github site:


  • Vilela, B., F. Villalobos, M.A. Rodriguez, and L.C. Terribile. 2014. Body size, extinction risk and knowledge bias in New World snakes. PLoS ONE doi:10.1371/journal.pone.0113429
  • Villalobos, F. and H.T. Arita. 2014. Morphological diversity at different spatial scales in a Neotropical bat assemblage. Oecologia 176, 557:568 doi: 10.1007/s00442-014-3039-y
  • Villalobos, F. and Rangel, T.F. 2014. Geographic patterns of biodiversity: macroecological approaches for a complex phenomenon. Contributed chapter for open-access book "Frontiers in Ecology, Evolution and Complexity" Edited by M. Benitez, O. Miramontes and A. Valiente-Banuet. Available at:
  • Arita, H.T., Vargas-Barón, J., and Villalobos, F. 2014. Latitudinal gradients of genus richness and endemism and the diversification of New World bats. Ecography 37, 1024:1033. doi: 10.1111/ecog.00720
  • Gouveia, S.F., Villalobos, F., Dobrovolski, R., Beltrão-Mendes, R., and Ferrari, S.F. 2014. Forest structure, not rainfall, drives global diversity of primates. Journal of Animal Ecology 83, 1523:1530. doi:10.1111/1365-2656.12241 Access the recommendation on F1000Prime
  • Bini, L.M., F. Villalobos and J.A.F. Diniz-Filho. 2014. Explorando patrones en rasgos macroecológicos utilizando regresión secuencial de autovectores filogenéticos. Ecosistemas 23(1), 21-26.
  • Villalobos, F. 2014. Linking diversity and distribution to understand biodiversity gradients and inform conservation assessments. Frontiers of Biogeography.
  • Villalobos, F., A. Lira-Noriega, J. Soberón and H.T. Arita. 2014. Co-diversity and co-distribution in phyllostomid bats: evaluating the relative roles of climate and niche conservatism. Basic and Applied Ecology 15, 85-91.


  • Villalobos, F., T.F. Rangel and J.A.F. Diniz-Filho. 2013. Phylogenetic fields of species: cross-species patterns of phylogenetic structure and geographic coexistence. Proceedings of the Royal Society B, Biological Sciences 280, 20122570.
  • Villalobos, F., R. Dobrovolski, D.B. Provete, and S.F. Gouveia. 2013. Is rich and rare the common share? Describing biodiversity patterns to inform conservation practices of South American anurans. PLoS ONE 8(2): e56073. doi:10.1371/journal.pone.0056073
  • Villalobos, F., A. Lira-Noriega, J. Soberón and H.T. Arita. 2013. Range-diversity plots for conservation planning: Using richness and rarity in priority setting, the case of Mexican avifauna. Biological Conservation 158: 313-320.


  • Provete, D.B., Villalobos, F. and Cianciaruso, M.V. 2012. Shadows of the past: paleo-reconstructions, phylogenies, and macroecological hypotheses. Update. Frontiers of Biogeography, the scientific journal of the International Biogeography Society. 4(2): 54-55.
  • Villalobos, F., L. Garrido-Olvera and P. Trejo-Barocio. 2012. Biogeography and ecology, are there signs of integration? Biogeografía: Bulletin of the Systematic and Evolutionary Biogeographical Association. 5: 3-7.


  • Villalobos, F. and O. Paknia. 2011. Biogeography and ecology: two lenses in one telescope. Symposium Summary. Frontiers of Biogeography, the scientific journal of the International Biogeography Society. 3(1): 5-7.
  • Barve, N., V. Barve, A. Jiménez-Valverde, A. Lira-Noriega, S.P. Maher, A.T. Peterson, J. Soberón, F. Villalobos. 2011. The crucial role of the accessible area in ecological niche modeling and species distribution modeling. Ecological Modelling. 222, 1810-1819.


  • Villalobos, F. and H.T. Arita. 2010. The Diversity Field of New World Leaf-Nosed Bats (Phyllostomidae). Global Ecology and Biogeography 19: 200-211.
  • Ortiz-Ramírez, D.R. and F. Villalobos. Wing morphology of four insectivore bats from Yucatan. Comparison between two methods to obtain morphological parameters. 2010. In Cervantes, F., Vargas, J. and Y. Hortelano (eds.) “60 años de la Colección Nacional de Mamíferos del Instituto de Biología, UNAM”


  • Jardim, L.L.C.Z, Bini, L.M., Diniz-Filho, J.A.F., and Villalobos, F. Challenging the Raunkiaeran shortfall and the consequences of using imputed databases. Available at bioRxiv
  • Velasco, J., Villalobos, F., Diniz-Filho, J.A.F., Poe, S., Flores-Villela, O. A comprehensive framework to evaluate the spatio-temporal dimensions of body size evolution in Anolis lizards. Submitted to Ecography
“...con frecuencia la ignorancia engendra más confianza que el conocimiento...”